<p>尝试一下:</p>
<pre><code>import re:
pat = re.compile(r'(\w{3})@')
txt = "(Esy@ESY15_g64743_DN3_SP7_c0:0.0726396855636,Aar@AA_maker7399_1:0.137507902808,((Spa@Tp2g18720:0.0318934795022,Cpl@CP2_g48793_DN3_SP8_c:0.0273465005242):9.05326020871e-05,(((Bst@Bostr_13083s0053_1:0.0332592496158,((Aly@AL8G21130_t1:0.0328569260951,Ath@AT5G48370_1:0.0391706378372):0.0205924636564,(Chi@CARHR183840_1:0.0954469923893,Cru@Carubv10026342m:0.0570981548016):0.00998579652059):0.0150356382287):0.0340484449097,(((Hco@scaff1034_g23864_DN3_SP8_c_TE35_CDS100:0.00823215335663,Hlo@DN13684_c0_g1_i1_p1:0.0085462978729):0.0144626717872,Hla@DN22821_c0_g1_i1_p1:0.0225079453622):0.0206478928557,Hse@DN23412_c0_g1_i3_p1:0.048590776459):0.0372829371381):0.00859075940423,(Esa@Thhalv10004228m:0.0378509854703,Aal@Aa_G102140_t1:0.0712272454125):1.00000050003e-06):0.00328120860999):0.0129090235079):0.0129090235079;"
pat.findall(t)
</code></pre>
<p>结果:</p>
<pre><code>['Esy', 'Aar', 'Spa', 'Cpl', 'Bst', 'Aly', 'Ath', 'Chi', 'Cru', 'Hco', 'Hlo', 'Hla', 'Hse', 'Esa', 'Aal']
</code></pre>
<p>如果您需要完整的结构,我们可以尝试移除不必要的部分:</p>
<pre><code>pat = re.compile(r'(@|:)[^/),]*')
pat.sub('',t).replace(',', ', ')
</code></pre>
<p>结果:</p>
<pre><code>'(Esy, Aar, ((Spa, Cpl), (((Bst, ((Aly, Ath), (Chi, Cru))), (((Hco, Hlo), Hla), Hse)), (Esa, Aal))))'
</code></pre>
<p><a href="https://regex101.com/r/KGQfRl/1" rel="nofollow noreferrer">Regex demo</a></p>